substrate -- (the substance that is acted upon by an enzyme or ferment)
substrate, substratum -- (a surface on which an organism grows or is attached; "the gardener talked about the proper substrate for acid-loving plants")
substrate, substratum -- (any stratum or layer lying underneath another)
substrate, substratum -- (an indigenous language that contributes features to the language of an invading people who impose their language on the indigenous population; "the Celtic languages of Britain are a substrate for English")